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The Ordovician Portal

Introduction

The Ordovician (/ɔːrdəˈvɪʃi.ən, -doʊ-, -ˈvɪʃən/ or-də-VISH-ee-ən, -⁠doh-, -⁠VISH-ən) is a geologic period and system, the second of six periods of the Paleozoic Era, and the second of twelve periods of the Phanerozoic Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 Ma (million years ago) to the start of the Silurian Period 443.8 Ma.

The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress.

Life continued to flourish during the Ordovician as it had in the earlier Cambrian Period, although the end of the period was marked by the Ordovician–Silurian extinction events. Invertebrates, namely molluscs and arthropods, dominated the oceans, with members of the latter group probably starting their establishment on land during this time, becoming fully established by the Devonian. The first land plants are known from this period. The Great Ordovician Biodiversification Event considerably increased the diversity of life. Fish, the world's first true vertebrates, continued to evolve, and those with jaws may have first appeared late in the period. About 100 times as many meteorites struck the Earth per year during the Ordovician compared with today in a period known as the Ordovician meteor event. It has been theorized that this increase in impacts may originate from a ring system that formed around Earth at the time. (Full article...)

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Selected article on the Ordovician world and its legacies

A life restoration of an ostracoderm.

A fish is any member of a paraphyletic group of organisms that consist of all gill-bearing aquatic craniate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups. Most fish are ectothermic ("cold-blooded"), allowing their body temperatures to vary as ambient temperatures change. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams to even hadal depths of the deepest oceans. At 32,000 species, fish exhibit greater species diversity than any other group of vertebrates. Because the term "fish" is defined negatively, and excludes the tetrapods (i.e., the amphibians, reptiles, birds and mammals) which descend from within the same ancestry, it is paraphyletic, and is not considered a proper grouping in systematic biology.

The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessednotochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. (see more...)

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Selected article on the Ordovician in human science, culture and economics

The history of paleontology traces the history of the effort to study the fossil record left behind by ancient life forms. Although fossils had been studied by scholars since ancient times, the nature of fossils and their relationship to life in the past became better understood during the 17th and 18th centuries. At the end of the 18th century the work of Georges Cuvier ended a long running debate about the reality of extinction and led to the emergence of paleontology as a scientific discipline.

The first half of the 19th century saw paleontological activity become increasingly well organized. This contributed to a rapid increase in knowledge about the history of life on Earth, and progress towards definition of the geologic time scale. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. After Charles Darwin published Origin of Species in 1859, much of the focus of paleontology shifted to understanding evolutionary paths.

The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection, as demonstrated by a series of important discoveries in China near the end of the 20th century. There was also a renewed interest in the Cambrian explosion that saw the development of the body plans of most animal phyla. (see more...)

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A fossil of the trilobite Pseudoasaphus praecurrens. This specimen dates back 460 – 468 Million years ago to the Darriwilian age of the Middle Ordovician epoch and was discovered near Saint Petersburg, Russia in the Koporka River deposits. It is 18.4 x 10 x 4.4 cm in size and has a mass of about 580g.
Photo credit: Didier Descouens

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Did you know?

...that eighty years on, scientists are still debating whether the Palæozoic fossils known as Chitinozoans (SEM image pictured) represent plants, animals or eggs?
... that small shells of Trigonoconcha are triangular?
... that a fossil of Concavodonta described in 1843 has been lost?
... that one species of the extinct bivalve Similodonta was found in 108.90 metres (357.3 ft) down a Welsh borehole?