Portal:Algae

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Introduction

A variety of algae growing on the sea bed in shallow waters

A variety of microscopic unicellular and colonial freshwater algae

Algae (UK: /ˈælɡiː/ AL-ghee, US: /ˈældʒiː/ AL-jee; sg.: alga /ˈælɡə/ AL-gə) is an informal term for any organisms of a large and diverse group of photosynthetic eukaryotes, which include species from multiple distinct clades. Such organisms range from unicellular microalgae such as Chlorella, Prototheca and the diatoms, to multicellular macroalgae such as the giant kelp, a large brown alga which may grow up to 50 metres (160 ft) in length. Most algae are aquatic organisms and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds. In contrast, the most complex freshwater forms are the Charophyta, a division of green algae which includes, for example, Spirogyra and stoneworts. Algae that are carried passively by water are plankton, specifically phytoplankton.

Algae constitute a polyphyletic group since they do not include a common ancestor, and although their chlorophyll-bearing plastids seem to have a single origin (from symbiogenesis with cyanobacteria), they were acquired in different ways. Green algae are a prominent examples of algae that have primary chloroplasts derived from endosymbiont cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from endosymbiotic red algae, which they acquired via phagocytosis. Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction via spores.

Algae lack the various structures that characterize plants (which evolved from freshwater green algae), such as the phyllids (leaf-like structures) and rhizoids of bryophytes (non-vascular plants), and the roots, leaves and other xylemic/phloemic organs found in tracheophytes (vascular plants). Most algae are autotrophic, although some are mixotrophic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy, myzotrophy or phagotrophy. Some unicellular species of green algae, many golden algae, euglenids, dinoflagellates, and other algae have become heterotrophs (also called colorless or apochlorotic algae), sometimes parasitic, relying entirely on external energy sources and have limited or no photosynthetic apparatus. Some other heterotrophic organisms, such as the apicomplexans, are also derived from cells whose ancestors possessed chlorophyllic plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria that produce oxygen as a byproduct of splitting water molecules, unlike other organisms that conduct anoxygenic photosynthesis such as purple and green sulfur bacteria. Fossilized filamentous algae from the Vindhya basin have been dated to 1.6 to 1.7 billion years ago. (Full article...)

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Selected general article

Pyrenoids are sub-cellular phase-separated micro-compartments found in chloroplasts of many algae, and in a single group of land plants, the hornworts. Pyrenoids are associated with the operation of a carbon-concentrating mechanism (CCM). Their main function is to act as centres of carbon dioxide (CO₂) fixation, by generating and maintaining a CO₂ rich environment around the photosynthetic enzyme ribulose-1,5-bisphosphate carboxylase/oxygenase (RuBisCO). Pyrenoids therefore seem to have a role analogous to that of carboxysomes in cyanobacteria.

Algae are restricted to aqueous environments, even in aquatic habitats, and this has implications for their ability to access CO₂ for photosynthesis. CO₂ diffuses 10,000 times slower in water than in air, and is also slow to equilibrate. The result of this is that water, as a medium, is often easily depleted of CO₂ and is slow to gain CO₂ from the air. Finally, CO₂ equilibrates with bicarbonate (HCO₃⁻) when dissolved in water, and does so on a pH-dependent basis. In sea water for example, the pH is such that dissolved inorganic carbon (DIC) is mainly found in the form of HCO₃⁻. The net result of this is a low concentration of free CO₂ that is barely sufficient for an algal RuBisCO to run at a quarter of its maximum velocity, and thus, CO₂ availability may sometimes represent a major limitation of algal photosynthesis. (Full article...)

Selected algae type

Examples of xanthophytes (repair the unequal flagella in the cells)

Yellow-green algae or the Xanthophyceae (xanthophytes) are an important group of heterokont algae. Most live in fresh water, but some are found in marine and soil habitats. They vary from single-celled flagellates to simple colonial and filamentous forms. Xanthophyte chloroplasts contain the photosynthetic pigments chlorophyll a, chlorophyll c, β-carotene, and the carotenoid diadinoxanthin. Unlike other Stramenopiles (heterokonts), their chloroplasts do not contain fucoxanthin, which accounts for their lighter colour. Their storage polysaccharide is chrysolaminarin. Xanthophyte cell walls are produced of cellulose and hemicellulose. They appear to be the closest relatives of the brown algae. (Full article...)

Selected images

Image 1Algae at Yellowstone Park
Image 2Stigeoclonium, a chlorophyte green alga genus
Image 3Eucheuma farming (underwater), Philippines
Image 4Aerial view of algae farm ponds for production of beta-carotin, Whyalla, South Australia.
Image 5Zygnema. 10× objective, 15× eyepiece; numbered ticks are 122 µM apart.
Image 6Micrasterias radiata / from Shishitsuka-Pond, Tsuchiura, Ibaraki Pref., Japan / Microscope: Leica DMRD (DIC)

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